Klug Research Group

Computational Biomechanics at UCLA






Reprints are available for personal use by request. E-mail klug@seas.ucla.edu


  • Missel AR, Bai M, Klug WS, Levine AJ, The Affine to Non-affine Transition in Anisotropic Semiflexible Networks.
  • Roos WH, Gibbons MM, Arkhipov A, Uetrecht C, Watts NR, Wingfield PT, Steven AC, Heck AJR, Schulten K, Klug WS, Wuite GJL, Squeezing protein shells: how continuum elastic models, moleculardynamics simulations and experiments coalesce at the nanoscale.


  • Morozov AY, Rudnick J, Bruinsma RF and Klug WS, Assembly and Disassembly of Deltahedral Viral Shells. In: Stockley, P. and Twarock, R., Emerging Topics in Physical Virology (London: Imperial College Press, 2010).
  • Hoshino T., Chow LA, Hsu JJ, Perlowski AA, Abedin M, Tobis J, Tintut Y, Mal AK, Klug WS, Demer LL, Mechanical stress analysis of a rigid inclusion in distensible material: a model of atherosclerotic calcification and plaque vulnerability., Am. J. Physiol.-Heart Circul. Physiol., 2009, 297 (2), H802-10.
  • Ursell TS, Klug WS, Phillips R, Morphology and interaction between lipid domains., Proc. Natl. Acad. Sci. U.S.A., 2009, 106 (32), 13301-6.
  • Lee HJ, Peterson EL, Phillips R, Klug WS, Wiggins PA, Membrane shape as a reporter for applied forces., Proc. Natl. Acad. Sci. U.S.A., 2008, 105 (49), 19253-7.
  • Wilkinson PR, Klug WS, Van Leer B, Gimzewski JK, Nanomechanical properties of piezoresistive cantilevers: Theory and experiment, J. Appl. Phys., 2008, 104 (10), -.
  • Gibbons MM, Klug WS, Influence of nonuniform geometry on nanoindentation of viral capsids, Biophys J, 2008, 95 (8), 3640-9.
  • Gibbons MM, Klug WS, Nonlinear finite-element analysis of nanoindentation of viral capsids, Phys Rev E Stat Nonlin Soft Matter Phys, 2007, 75 (3 Pt 1), 031901.
  • Klug, W.S., Bruinsma, R.F., Michel, J.-P., Knobler, C.M., Ivanovska, I.L., Schmidt, C.F., and Wuite, G.J.L., Failure of Viral Shells, Phys Rev Lett, 97, 228101 (2006). [doi]
  • F. Feng and W.S. Klug, Finite Element Modeling of Lipid Bilayer Membranes, Journal of Computational Physics, 20(1), 394-408 (2006). [doi]
  • J.-P. Michel, I.L. Ivanovska, M.M. Gibbons, W.S. Klug, C.M. Knobler, G.J.L. Wuite, and C.F. Schmidt, Nanoindentation Studies of Full and Empty Viral Capsids and the Effects of Capsid Protein Mutations on Elasticity and Strength, Proceedings of the National Academy of Sciences, 103(16):6184 - 6189 (2006). [doi]
  • J. Reed, P. Wilkinson, J. Schmit, W. Klug, and J.K. Gimzewski, Observation of Nanoscale Dynamics in Cantilever Sensor Arrays, Nanotechnology, 17:3873-3879 (2006). [doi]
  • W.S. Klug, M.T. Feldmann, M. Ortiz, Three-dimensional director-field predictions of viral DNA packing arrangements, Computational Mechanics, 35(2), 146-152, 2005. [doi]
  • W. S. Klug, M. Ortiz, A Director Field Model of DNA Packaging in Viral Capsids, Journal of Mechanics and Physics of Solids, 51(10):1815-1847, 2003. [doi]
  • A. Mota, W. Klug, A. Pandolfi, M. Ortiz, Finite Element Simulation of Firearm Injury to the Human Cranium, Computational Mechanics, 31(1-2): 115-121, 2003. [doi]

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